Fender0740 wrote:Now sorry if this is a stupid question but what do they contribute towards in tone?
Not stupid at all. Everything starts with the transformers. The PT transforms AC power to higher voltage AC where it is then converted to DC by the rectifier. At the end of the chain, the OT transforms DC power back to AC to power the speaker. Everything in the signal path has a sound; resistors, capacitors, the tubes themselves, the wire. Their physical characteristics and materials behave in a certain way and influence the shape of the wave moving through them. The sum of these parts and even the placement of the components and wiring (lead dress) create the tone of an amplifier. Great, great debate and partisanship exists over this.
For the manufacturer's part, the quality of materials, precision (reproducibility from part to part), design, all play a role in how the part ends up performing. The OT is especially important as we are concerned with how it performs over a range of frequencies. Valve Juniors sound pretty awful out of the box due to their weak, anemic OT, low quality parts. But they are built to a price point; their job isn't to sound good, it's to be inexpensive. Replace the OT with something of better design, quality and bandwidth, and whoa, there's a good amp in there.
You can always hear the difference with a hand wired amp made from quality components. The difference however is philosophical as tone is in the ear of the beerholder. There is no right or wrong answer, only what you like.